Many photos of the blesbok (Damaliscus pygargus phillipsi) show individuals in which
This spreading of the facial bleeze - which I have not seen mentioned in the literature - may possibly be a sign of age (exceeding ten years). This is indicated by the fact that many of the photos concerned are of captive specimens.
The ultimate result is seen in https://www.inaturalist.org/observations/70027286.
Most of the photos below refer to females, but males also show this pattern (e.g. https://www.inaturalist.org/observations/141372682 and https://www.inaturalist.org/observations/32952125 and https://www.flickr.com/photos/holtphoto/11667853505/in/photolist-iM3Nm2-6TeDse-29Yu8yw-GQHdeu-QzVdjX-e6MgFY-2gjCi6T-H9VkFH-5oDPzj-ASgX6y-H7BJzh-pi4vHm-eYXy2U-5f8mw4-bFbjtX-YaJrFP-pHAQwC-24MnFrZ-71RcCB-csqwjA-3yg1Bw-4GLgxP-6UtPYS-5QZpnc-6DNMdK-37BsTy-dM3YJg-2nbqYFY-275Jw9t-C55EVf-apLdKb-rDKi6z-qtrvU-2nidniQ-o3mr2c-2jjUxYq-2hsDGV3-2hsGrgp-2iGfCyR-HfX4Cz-qtwsg8-ApjDMc-2jfZf6T-zEj283-qpGt6C-bt1qDP-dcWPMW-nQmxok-qFeiSy-2bW3UGB).
We now know of at least three aspects of the colouration of the blesbok that show particular depigmentation, compared to the bontebok (Damaliscus pygargus pygargus), viz.
The pattern on the cheeks, described here, occurs also in the bontebok (https://www.inaturalist.org/observations/27251980). However, it is faint and uncommon in that subspecies.
https://www.dreamstime.com/blesbok-lying-grass-blesbok-image265633507
https://www.inaturalist.org/observations/128960710
https://www.inaturalist.org/observations/77675104
https://www.shutterstock.com/it/image-photo/antelope-wild-blesbok-blesbuck-lying-down-2223739221
https://www.euro-t-guide.com/See_Photo/Denmark/JUT-M_Grenaa/Ree_Park_Ebeltoft-2008-14.jpg
https://depositphotos.com/635653818/stock-photo-blesbok-blesbuck-damaliscus-pygargus-phillipsi.html
https://www.shutterstock.com/it/image-photo/blesbok-latin-name-damaliskus-dorcas-phillipsi-360070883
https://www.shutterstock.com/it/image-photo/blesbok-latin-name-damaliskus-dorcas-phillipsi-290826794
https://www.shutterstock.com/it/image-photo/blesbuck-grazing-on-green-grass-1275827710
https://dewetswild.com/2016/06/23/blesbok/#jp-carousel-52456
https://www.inaturalist.org/observations/35368437
https://www.inaturalist.org/observations/124183870
https://www.dreamstime.com/stock-image-blesbok-close-up-antelope-eating-image30613991 and https://www.dreamstime.com/stock-images-blesbok-close-up-antelope-image30613954
Please note the extremely long horns, indicating advanced age, in the central figure in https://www.inaturalist.org/observations/34887688
Also see https://www.inaturalist.org/journal/milewski/75748-adaptive-colouration-in-the-blesbok-damaliscus-pygargus-phillipsi# and https://www.inaturalist.org/posts/78685-the-overlooked-complexity-of-facial-markings-in-genus-damaliscus#.
For an index to my many Posts about the genus Damaliscus, please see https://www.inaturalist.org/journal/milewski/78238-an-index-to-my-posts-on-genus-damaliscus#.
Comments
The following adult male individual, which is territorial judging from the glandular staining on the facial bleeze, seems to show the beginning of an expansion of pale on the sides of the face:
https://www.inaturalist.org/observations/21052176
https://www.dreamstime.com/stock-photo-blesbok-antelopes-damaliscus-pygargus-isolated-white-background-image59171426
https://www.farmersweekly.co.za/animals/game-and-wildlife/a-promising-new-colour-variant-for-game-farmers/
http://eswatininaturereserves.com/mammals/redlist/2016MammalRedListBlesbok-Damaliscus-pygargus-phillipsi_LC.pdf
https://www.researchgate.net/publication/312232146_Blesbok_Damaliscus_pygargus_phillipsi_Bontebok_Damaliscus_pygargus_pygargus
https://link.springer.com/article/10.1007/s12686-018-0983-6
https://onlinelibrary.wiley.com/doi/full/10.1002/ece3.2595
https://stock.adobe.com/search?filters%5Bcontent_type%3Aphoto%5D=1&filters%5Bcontent_type%3Aillustration%5D=1&filters%5Bcontent_type%3Azip_vector%5D=1&filters%5Bcontent_type%3Avideo%5D=1&filters%5Bcontent_type%3Atemplate%5D=1&filters%5Bcontent_type%3A3d%5D=1&filters%5Bcontent_type%3Aaudio%5D=0&filters%5Binclude_stock_enterprise%5D=0&filters%5Bis_editorial%5D=0&filters%5Bfree_collection%5D=0&filters%5Bcontent_type%3Aimage%5D=1&k=damaliscus&order=relevance&safe_search=1&limit=100&search_page=14&search_type=pagination&get_facets=0&asset_id=86951676
https://stock.adobe.com/search?filters%5Bcontent_type%3Aphoto%5D=1&filters%5Bcontent_type%3Aillustration%5D=1&filters%5Bcontent_type%3Azip_vector%5D=1&filters%5Bcontent_type%3Avideo%5D=1&filters%5Bcontent_type%3Atemplate%5D=1&filters%5Bcontent_type%3A3d%5D=1&filters%5Bcontent_type%3Aaudio%5D=0&filters%5Binclude_stock_enterprise%5D=0&filters%5Bis_editorial%5D=0&filters%5Bfree_collection%5D=0&filters%5Bcontent_type%3Aimage%5D=1&k=damaliscus&order=relevance&safe_search=1&limit=100&search_page=14&search_type=pagination&get_facets=0&asset_id=262944266
https://stock.adobe.com/search?filters%5Bcontent_type%3Aphoto%5D=1&filters%5Bcontent_type%3Aillustration%5D=1&filters%5Bcontent_type%3Azip_vector%5D=1&filters%5Bcontent_type%3Avideo%5D=1&filters%5Bcontent_type%3Atemplate%5D=1&filters%5Bcontent_type%3A3d%5D=1&filters%5Bcontent_type%3Aaudio%5D=0&filters%5Binclude_stock_enterprise%5D=0&filters%5Bis_editorial%5D=0&filters%5Bfree_collection%5D=0&filters%5Bcontent_type%3Aimage%5D=1&k=damaliscus&order=relevance&safe_search=1&limit=100&search_page=15&search_type=pagination&get_facets=0&asset_id=221411745
https://www.inaturalist.org/observations/124183870
Extension of whitish on to the forehead:
https://stock.adobe.com/search/images?filters%5Bcontent_type%3Aphoto%5D=1&filters%5Bcontent_type%3Aillustration%5D=1&filters%5Bcontent_type%3Azip_vector%5D=1&filters%5Bcontent_type%3Avideo%5D=1&filters%5Bcontent_type%3Atemplate%5D=1&filters%5Bcontent_type%3A3d%5D=1&filters%5Bcontent_type%3Aaudio%5D=0&filters%5Binclude_stock_enterprise%5D=0&filters%5Bis_editorial%5D=0&filters%5Bfree_collection%5D=0&filters%5Bcontent_type%3Aimage%5D=1&k=damaliscus&order=relevance&safe_search=1&limit=100&search_page=8&search_type=pagination&get_facets=0&asset_id=547249274
https://zooinstitutes.com/animals/blesbok-566/
https://www.alamy.com/stock-image-herd-of-blesbok-antelope-running-across-the-dusty-ground-in-limpopo-163785330.html?imageid=30F56100-3C57-4059-83B8-EC293683DAE3&p=218068&pn=1&searchId=1acbb4918a3fb248f068c8228e9df229&searchtype=0
The following is possibly the clearest illustration I have ever seen of multi-facetted extension of the facial bleeze in adult males of the blesbok:
https://www.inaturalist.org/observations/175192441
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